Optimus explains that the Proudstar, captained by Nova Prime, was a ship meant to convert uninhabited planets into ones suitable to support Cybertronian life, and that its Ferrotaxis supercomputer contains a record of Cybertron's history and culture which had been lost during the Great War. Entering the ship, the Autobots fight off Soundwave and attempt to access the plasma core powering the Proudstar and the Insecticons, but the security system ejects the core from the ship. Emerging, the Autobots learn the core's location from Thundercracker and follow the trail. While they are distracted by fighting Blitzwing, Starscream arrives to take the core back to the Decepticons, but loses control of it due to a storm and crash-lands. The Autobots beat Motormaster to the core, but find it damaged and at risk of explosion. Before they can shut it down, the rest of the Stunticons arrive and form Menasor. The combiner throws the core to Megatron, who takes it to Cybertron to be properly repaired.
The Autobots follow the Decepticons through the Space Bridge, but are delayed by Starscream and Shockwave, arriving too late to catch Megatron. Conversing with Shockwave, Optimus discovers that during the Proudstar 's voyage, Nova Prime and the crew were corrupted by Unicron and had begun to cyberform inhabited planets as well, planning to do the same to Earth before they crashed. Returning to the Proudstar, the Autobots try once again to disable the core, but it ruptures and becomes inaccessible, leaving the Ferrotaxis, now in Megatron's possession, as the only way to stop the cyberforming process. The group head for the Proudstar 's control room, fighting off the Decepticon forces as they ascend. Megatron escapes, ordering Devastator and Menasor to finish them, but the Autobots defeat them and follow Megatron, fighting him for the computer.
As he is defeated, Megatron fires the Ferrotaxis from his cannon into orbit, and Optimus rides an Insecticon into space after it, with Megatron in pursuit. The two engage in a final battle above the Earth, with Optimus emerging victorious, but cannot shut down the Insecticons due to Megatron's tampering. Left with no choice, Optimus destroys the Ferrotaxis, stopping the cyberforming and preventing the plasma core's detonation. As he falls back to Earth, Optimus declares Earth is now their true home, and they must protect it at all costs. Later, with Defensor and Superion's support, Optimus commissions plans for the Optimus Maximus project.
In a post-credits scene, aboard the Proudstar, Nova Prime is seen awakening from stasis.
Gameplay
Transformers: Devastation is an action game similar to PlatinumGames's other titles. Battling against Decepticon forces within large 3D environments, players are able to control one of five Autobots: Optimus Prime, Bumblebee, Sideswipe, Wheeljack, and Grimlock. Gameplay includes fast-paced combos as well as the ability to switch between vehicle and robot modes. A customization system allows players to find and upgrade various weapons for each character, as well as equip T.E.C.H. that grant passive boosts to characters' stats.
The game is part of the Transformers: Generations line, though its visual design is most strongly inspired by the original Transformers TV series. The game's story is written by former Transformers comic writer and editor Andy Schmidt. Several voice actors reprise their roles from the original show, including Peter Cullen (Optimus Prime), Dan Gilvezan (Bumblebee), Michael Bell (Sideswipe and Scrapper), Gregg Berger (Grimlock and Long Haul), and Frank Welker (Megatron and Soundwave). Original music was composed by Vince DiCola (the composer of The Transformers: The Movie and Angry Birds Transformers) and Kenny Meriedeth.
A downloadable content bundle containing character skins based on Nemesis Prime, Red Alert, and Goldfire, along with three additional weapons, was offered as a pre-order incentive. This bundle was later made available for separate purchase
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